Biological and Environmental Sciences M.S., Ph.D.

Specialization: Integrative and Evolutionary Biology (IEB)

This graduate group focuses on the diversity of form and function of organisms from evolutionary and physiological perspectives, as well as the application of these approaches to health, agriculture, and the environment. Specific faculty research interests are diverse and include marine biology, animal science (including reproduction, nutrition, management, and health), aquaculture (including ecology, physiology, nutrition, and health), cellular and behavioral neurobiology (including sensory biology and neuroethology), evolutionary biology, genomics (comparative, evolutionary, and marine), morphology and development (including functional morphology, biomechanics, and evolutionary developmental biology), paleontology, physiology and pathology (including environmental, stress, reproductive, and comparative physiology, endocrinology, aquatic pathology), plant biology, and human health.

Prospective Students

A strong advisor-student relationship is central to the IEB program. Prospective students should contact faculty directly to explore mutual interests before applying. Program flexibility encourages our students to work across traditional departmental boundaries and often seek interdisciplinary collaborations.

What should I submit with my online application?

All applications must be submitted online using the ApplyYourself system. EES application deadlines are outlined in the FAQs section. For access to the system and detailed instructions on what must be submitted, please visit the URI Graduate School web pages. In addition to the graduate school submission requirements, please make sure the following are also included in your application materials submitted on ApplyYourself:

  1. At the beginning of your personal statement, list one or more potential advisors in bold-face type.
  2. Arrange for submission of two letters of recommendation.
  3. Submit a scan of your GRE scores, uploaded as “supplemental materials” (in addition to requesting that ETS send a copy directly to URI).
  4. Consider submitting any additional materials that provide direct evidence of relevant experience or writing ability, such as published papers, research posters, or reports. Please upload any such documents as “supplemental materials”.


Specialization group coordinator: Steve Irvine,
Associate Professor of Biological Sciences
College of the Environment and Life Sciences
Phone: 401-874-9204
Email: steven.irvine@uri.edu