Course Descriptions

BIO 101

Principles of Biology I

(3 crs.) Chemistry, structure, metabolism, and reproduction of cells. Principles of genetics. Structure, development, and physiology of animals. Survey of the animal kingdom. (Lec. 3/Online) Pre: Credit or concurrent enrollment in BIO 103. (A1)

BIO 101

Principles of Biology I

(3 crs.) Chemistry, structure, metabolism, and reproduction of cells. Principles of genetics. Structure, development, and physiology of animals. Survey of the animal kingdom. (Lec. 3/Online) Pre: Credit or concurrent enrollment in BIO 103. Not open to students with credit in BIO 110. (A1)

BIO 101H

Honors Section of BIO 101: Principles of Biology I

(3 crs.) Honors Section of BIO 101: Chemistry, structure, metabolism, and reproduction of cells. Principles of genetics. Structure, development, and physiology of animals. Survey of the animal kingdom. (Lec. 3) Pre: Credit or concurrent enrollment in BIO 103 and a 3.4 overall GPA. (A1)

BIO 101H

Honors Section of BIO 101: Principles of Biology I

(3 crs.) Honors Section of BIO 101: Chemistry, structure, metabolism, and reproduction of cells. Principles of genetics. Structure, development, and physiology of animals. Survey of the animal kingdom. (Lec. 3) Pre: Credit or concurrent enrollment in BIO 103 and a 3.4 overall GPA. Not open to students with credit in BIO 110. (A1)

BIO 102

Principles of Biology II

(3 crs.) Structure, physiology, and reproduction of plants. Diversity of plants, fungi, and algae. Principles of ecology and evolution. (Online) Pre: BIO 101 or 110, 103, and credit or concurrent enrollment in BIO 104. (A1)

BIO 103

Principles of Biology Laboratory I

(1 cr.) Selected laboratory exercises to accompany BIO 101. (Lab. 2) Pre: credit or concurrent enrollment in BIO 101. (A1)

BIO 103

Principles of Biology Laboratory I

(1 cr.) Selected laboratory exercises to accompany BIO 101. (Lab. 2) Pre: credit or concurrent enrollment in BIO 101 or 110. (A1)

BIO 104

Principles of Biology Laboratory II

(1 cr.) Selected laboratory exercises to accompany BIO 102. (Lab. 2) Pre: BIO 101 or 110, 103 and credit or concurrent enrollment in 102. (A1)

BIO 110

Fundamentals of Biology

(3 crs.) Fundamental biological concepts including the chemical and cellular basis of life, genetics, evolution, and organismal form and function. Intended for science majors who require one semester of biology. (Lec. 3) Pre: Credit or concurrent enrollment in BIO 103. Not open to students with credit in BIO 101. (A1)

BIO 122X

Biology and Behavior of Sharks and Whales

(3 crs.) Cross-listed as (BIO) AVS 122X. Develop foundational skills in science by conducting research on sharks and whales. Three weekend field experiences are required. (Lec. 2, Lab. 1) Pre: First semester freshmen only.

BIO 130

Topics In Marine Biology

(1 cr.) Current and classical issues considered in small classes. Designed for students interested in marine biology. (Seminar) Pre: Limited to marine biology majors. Required of all freshmen marine biology majors and students entering the major with fewer than 24 credits. May not be repeated.

BIO 181G

The Information Age: From Politics to Medicine

(3 crs.) Cross-listed as (BIO), DSP 181G. How big data affects our society, from advertising to politics to medicine. (Lec. 3/Online) Not for major credit for B.S. Biological Sciences or B.A. Biology. (A1) (GC)

BIO 191X

First-Year Research in Biology

(2 crs.) Introduces students to research in biology through hands-on projects. Helps students make a plan to reach individual goals and develop research skills in biology they can use to accomplish these goals. (Seminar) Pre: Completed or concurrent enrollment in BIO 101, freshman standing, or by permission of instructor. S/U credit.

BIO 201

General Animal Physiology

(4 crs.) The study of animal physiology (respiration, bioenergetics, locomotion, circulation, osmoregulation, hormones, nervous system, sensory organs, reproduction, immune function) with a consideration of environmental challenges and evolutionary adaptations. (Lec. 3, Lab. 1) Pre: BIO 101/103 or 110/103, and 102/104, and CHM 101, or permission of instructor.

BIO 220

Fundamentals of Human Anatomy and Physiology I

(3 crs.) Explores the integration of the structures and functions of the human organism through a systems approach; including the biochemical properties of the cell, integumentary, muscular, skeletal, and nervous systems. (Lec. 3/Online) Pre: Credit or concurrent enrollment in BIO 221.

BIO 221

Fundamentals of Human Anatomy and Physiology I Laboratory

(1 cr.) Fundamental structure and function of organ systems of the human body explored through models, dissections, and experiments. Specific topics include: tissues, integumentary system, muscular system, skeletal system, and nervous system. (Lab. 1) Pre: Credit or concurrent enrollment in BIO 220.

BIO 222

Fundamentals of Human Anatomy and Physiology II

(3 crs.) Explores the integration of the structures and functions of the human organism through a systems approach; including senses, endocrine, cardiovascular, lymphatic, immune, respiratory, digestive, urinary, and reproductive. (Lec. 3/Online) Pre: BIO 220 and 221, and credit or concurrent enrollment in BIO 223.

BIO 223

Fundamentals of Human Anatomy and Physiology II Laboratory

(1 cr.) Fundamental structure and function of organ systems of the human body explored through models, dissections, and experiments. Specific topics include: special senses, endocrine, circulatory, respiratory, digestive, urinary, and reproductive systems. (Lab. 1) Pre: BIO 220 and BIO 221, and credit or concurrent enrollment in BIO 222.

BIO 228

Seminar in Biological Sciences

(1 cr.) Lectures by local scientists on recent and ongoing basic research in biology, especially on-campus studies. Follow-up discussions will support student understanding and prepare students for independent research. (Seminar) Pre: Credit or concurrent registration in BIO 102. S/U only.

BIO 230G

Communicating Science through Data Visualization

(3 crs.) Exploring how scientists and communicators can use data visualization to communicate complex concepts simply and effectively to wide and varied audiences. (Lec. 2, Online) Pre: Students must be of sophomore standing. (C1) (GC)

BIO 242

Introductory Human Physiology

(3 crs.) Functions of the organ systems of the human body and their coordination in the whole human organism. Attention is given to the needs of students preparing for health-related professions. (Lec. 3) Pre: BIO 121.

BIO 256G

Risking our Reefs: Human Impacts on Ecosystem Builders

(3 crs.) Examination of human impacts on coral and oyster reef ecosystems and the ecological, economic, and ethical considerations of reef use and restoration. (Lec. 3) Pre: BIO 262 or NRS 223. (A1) (C2) (GC)

BIO 262

Introductory Ecology

(4 crs.) Structure and function of ecosystems, limiting factors, population dynamics, population interactions, and community relationships. Selected habitats and general ecological effects of humans. (Lec. 3, Rec. 1) Pre: BIO 101 or 110, 102 or equivalent.

BIO 263

Introduction to Ecological Data Analysis

(1 cr.) Learn to analyze ecological data.Gain quantitative skills, reinforce ecological concepts, and learn to integrate concepts to answer biological questions. (Lec. 1) Pre: credit or concurrent registration in BIO 262.

BIO 263

Introduction to Ecological Data Analysis

(1 cr.) Cross-listed (BIO) NRS 263. Learn to analyze ecological data. Gain quantitative skills, reinforce ecological concepts, and learn to integrate concepts to answer biological questions. (Lec. 1) Pre: credit or concurrent registration in BIO 262.

BIO 272

Introduction to Evolution

(4 crs.) Cross-listed as (BIO), GEO 272. Introduction to evolution as the unifying thread in the biosphere. Processes and patterns discussed, including microevolution and macroevolution. Social impact of evolution discussed from a biological perspective. Pre: GEO 102 or one semester of biological sciences, or permission of instructors.

BIO 282G

Sapiens: The Changing Nature of Human Evolution

(3 crs.) Cross-listed with (BIO), APG 282G. Study of human origins and history in order to understand Homo sapiens as a significant cause of evolutionary change, including an in-depth description of our widespread influence on Earth's systems. (Lec. 3) Pre: Sophomore or higher standing. (A1) (C2) (GC)

BIO 286

Humans, Insects, and Disease

(3 crs.) Cross-listed as (BIO), ENT 286. Role of insects, ticks, and mites as vectors and as direct agents of diseases in humans; factors affecting the spread of these diseases and their role in our cultural development. (Lec. 3) Not for major credit for B.S. in biological sciences.

BIO 290X

Research in Evolutionary Biology

(3 crs.) Conduct novel research in evolutionary biology. Learn how we determine how species are related using genetic data. (Lec. 1, Practicum 2) Pre: BIO 102 and 104.

BIO 291

Laboratory Research Skills

(1 cr.) Cross-listing as (AFS), AVS, BIO 291. Overview of the general operations of a scientific laboratory, commonly used scientific infrastructure, equipment and skills to prepare students for working in professional laboratories. (Lab. 1, Online) Pre: Sophomore standing; permission of instructor. The course is not for lab credit for B.S. Biological Sciences & B.S. Marine Biology.

BIO 300

Physiology of Exercise

(3 crs.) Cross-listed as (KIN), BIO 300. Applied human physiology, with applications to physical activity, exercise, and sport. Particular attention to acute and chronic adjustments of the circulatory, respiratory, metabolic, and muscular systems with exercise. (Lec. 3/Online) Pre: BIO 121 or BIO 220 and BIO 221 and BIO 222 and BIO 223 or permission of instructor. Open to Kinesiology and Biology majors only.

BIO 301

Physiology of Exercise Laboratory

(1 cr.) Cross-listed as (KIN), BIO 301. Student participation in laboratory sessions designed to understand the physiology of exercise relating to body composition, EKG, pulmonary, and metabolic functions. (Lab. 2) Pre: BIO 220, BIO 221, BIO 222 and BIO 223, completion of or concurrent enrollment in KIN (BIO) 300 or permission of instructor. Open to Kinesiology and Biology majors only.

BIO 302

Animal Development

(4 crs.) Survey of the patterns and mechanisms of animal development, including the molecular genetic control of development, medical developmental biology and evolution of development. (Lec. 3, Lab 3) Pre: BIO 101 or 110, 102, and two additional semesters of biological sciences; genetics recommended.

BIO 308

The Invisible Living Ocean

(3 crs.) The goal of this course is to explore the major groups of marine organisms that are mostly invisible to the naked eye, which fuel food webs, drive global biogeochemical cycles and affect climate. (Lec. 3) Pre: BIO 101 or 110, and BIO 102, or permission of instructor.

BIO 310

Bermuda Marine Biodiversity

(2 crs.) Based at the Bermuda Institute for Ocean Sciences, this course will provide students with experience in biodiversity assessment in the field, where we will blend fieldwork, lectures and laboratory manipulation. (Lab. 6) Pre: concurrent enrollment in BIO308.

BIO 311

Plant Structure and Development

(4 crs.) Structure of vascular plant cells, tissues and organs; cellular and molecular mechanisms controlling developmental processes including cell division, leaf initiation, epidermal patterning and vascular differentiation. (Lec. 3, Lab. 3) Pre: BIO 102 or permission of instructor.

BIO 320

Introduction to Computational Biology

(3 crs.) Cross-listed as (CMB), BIO 320. Introduction to the current topics of computational biology. Students will obtain hands-on experiences in navigating biological databases and analyzing biological data. (Lec. 3) Pre: CMB 201 or CMB 211.

BIO 321

Plant Diversity

(4 crs.) Representative forms of prokaryotes, algae, fungi, bryophytes, and vascular plants with emphasis on evolution, ecology, and life cycle. (Lec. 3, Lab. 3) Pre: BIO 102 or permission of instructor.

BIO 323

Field Botany and Taxonomy

(4 crs.) Cross-list with (NRS), BIO 323. Collection, identification, and study of vascular flora of Rhode Island, including use of manuals and herbarium specimens. Field trips throughout Rhode Island. Discussion of principles, methods, and data used in classification. (Lab. 2, Online) Pre: BIO 102.

BIO 331

Parasitology

(3 crs.) An introduction to the biology of parasitic organisms, including life cycles, infection pathways, and impacts on hosts. Topics are explored via lecture, laboratory exercises, literature review, and discussion groups. (Online) Pre: BIO 101 and BIO 102.

BIO 332

Plant Pathology: Introduction to Plant Diseases

(4 crs.) Cross-listed as (BIO), PLS 332. Nature, cause, and control of plant diseases. Use of basic techniques for identification of major types of plant diseases and their causal agents. (Lec. 4) Pre: BIO 102 or PLS 150 or permission of instructor.

BIO 341

Principles of Cell Biology

(3 crs.) Cross-list as (BIO), CMB 341. An introduction to the structure and organization of eukaryotic cells. Topics include membranes and organelles, gene expression, protein synthesis and secretion, energy utilization, the cytoskeleton, and signal transduction. (Lec. 3) Pre: one semester of biological sciences and one semester of organic chemistry.

BIO 345

Marine Environmental Physiology

(4 crs.) The physiological basis of adaptation to the marine environment. Physiological methods used to study marine organisms (Lec. 3, Lab. 1) Pre: BIO 260 or 360 or permission of instructor.

BIO 346

Plant Physiology

(3 crs.) Development and function of vascular plants, including energy and nutrient assimilation, growth, reproduction, and interactions with other organisms and the physical environment. (Lec. 3) Pre: BIO 102, one semester of chemistry, or permission of instructor.

BIO 350

Field Entomology and Taxonomy

(4 crs.) Cross-listed as (ENT), BIO, NRS 350. Collection, identification and study of the common families of insects, including use of keys and teaching collection specimens. Field trips throughout Rhode Island. Discussion of insect classification. (Lec. 3, Lab. 1) Pre: BIO 102 or permission of instructor.

BIO 352

General Genetics

(4 crs.) Cross-listed as (BIO), CMB 352. Introduction to basic genetic principles and concepts leading to an understanding of genes, heredity, and the nature of inherited variation. Applications and implications for animals, plants, fungi and bacteria. (Lec. 3, Rec. 1) Pre: BIO 101 or 110, and BIO 102.

BIO 353

Genetics Laboratory

(1 cr.) Cross-listed as (CMB), BIO 353. Basic principles and concepts of genetics demonstrated with microorganisms, plants, and animals. (Lab. 2) Pre: credit or concurrent enrollment in CMB/BIO 352.

BIO 354

Invertebrate Zoology

(4 crs.) Study of the origin and evolutionary relationship of the invertebrate animals. Emphasis on marine forms. Laboratory sessions include comparative study of selected examples and field trips to local environments. (Lec. 3, Lab. 1) Pre: BIO 101 or 110, 102, 103, and 104.

BIO 355

Marine Invertebrates of Southern New England

(3 crs.) Collection and identification of marine invertebrates of southern New England. Emphasis on field work and laboratory studies. Students collection will incorporate video photography. (Lab. 6) Pre: BIO 101 or 110, and 102 or permission of instructor.

BIO 360

Marine Biology

(4 crs.) The nature of plants and animals of the sea. Diversity of species and adaptations to habitats from the sea surface to the depths of the ocean. (Lec. 3, Lab. 3) Pre: BIO 101 or 110, and 102.

BIO 365

Biology of Algae

(4 crs.) Taxonomy, morphology, and evolution of all major algal divisions. Laboratory/field component focuses upon taxonomic identification of both live and preserved microscopic and macroscopic algal species. (Lec. 3, Lab. 3) Pre: BIO 102.

BIO 366

Vertebrate Biology

(3 crs.) Life histories, adaptations, ecology, classifications, and distribution of vertebrate animals. Laboratory and extensive field work on local vertebrates. (Lec. 2, Lab 3) Pre: BIO 262 recommended.

BIO 372

Introductory Endocrinology

(3 crs.) Cross-listed (AVS), BIO 372. Morphology and physiology of endocrine glands. Roles of hormones in regulation of body processes. Discussion of all endocrine organs and relationship of endocrine and nervous systems. (Lec. 3) Pre: BIO 101 or BIO 110, sophomore standing, or permission of instructor.

BIO 385

Introductory Entomology

(3 crs.) Cross-listed as (ENT), BIO 385. Introduction to the diverse components of entomology, emphasizing basic principles of insect morphology, physiology, behavior, and ecology. Current topics in insect biodiversity and management strategies. (Lec. 3) Pre: BIO 102 and BIO 101 or 110,, or permission of instructor.

BIO 388

Biology of Bees and Pollination Ecology

(3 crs.) Cross-listed as (ENT), BIO, NRS 388. Learn the biology, behavior and pollination services of bees. Learn how to identify bees to family level. Understand the basics of honey bee management. Learn various types of pollination. (Lec. 3) Pre: BIO 101 or 110, and 102, or permission of instructor.

BIO 396

Biology And Society

(3 crs.) A seminar course dealing with the impact of biological discoveries on societal questions and with the social influences that affect biological discovery. Discussion of original papers, magazines, newspaper articles, and books about various discoveries. (Seminar) Pre: Pre: junior or senior standing majoring in Biological Sciences (BS), Biology (BA) or Marine Biology (BS), or permission of instructor. (D1) (B4)

BIO 404

(304) Comparative Vertebrate Anatomy

(4 crs.) Anatomy of chordates emphasizing functional and evolutionary diversity. Lecture focuses on morphological variation and evolution including study of primary literature. Laboratory focuses on comparative anatomy through dissections and models. (Lec. 3, Lab. 3) Pre: BIO 101 or 110, and 102 and junior standing.

BIO 412

Evolution and Diversity of Fishes

(4 crs.) Origin, evolution and diversification of fishes, their phylogenetic relationships, and morphological, physiological, ecological, and behavioral adaptations in marine and freshwater habitats. (Lec. 3, Lab. 3) Pre: BIO 101 or 110, and 102, or permission of instructor. Not for graduate credit.

BIO 416

Intertidal Ecology

(4 crs.) The ecology of intertidal shores within New England, with a focus on common marine macroalgae. Includes field surveys, specimen identification, and an emphasis on research planning and hypothesis testing. (Lec. 3, Lab. 1) Pre: BIO 101, 102 and BIO 262 or BIO 360 (or permission from instructor).

BIO 417

Herpetology

(4 crs.) Cross-listed as (NRS), BIO 417. Introduces students to the biology, ecology, conservation, and management of reptiles and amphibians, including global perspectives, and field methods for studying amphibians and reptiles of northeastern North America. (Lec. 2, Lab. 4) Pre: BIO 101/103 or 110/103, and 102/104; and NRS 223 or BIO 262, and permission of instructor. Not for graduate credit.

BIO 419

Field Experience in Herpetology

(1 cr.) Cross-listed as (NRS), BIO 419. Capstone field trip in herpetology to region with higher amphibian and reptile diversity, such as Appalachia, to hone skills in identification, broaden understanding of ecology, and apply field research methods. (Practicum) Pre: concurrent enrollment in or credit for NRS/BIO 417, and permission of instructor. S/U only. Not for graduate credit.

BIO 422

Biology of Sharks and Their Relatives

(3 crs.) Survey of sharks, skates and rays including their classification, evolutionary history, physiology, ecology and interactions with humans. (Lec. 3) Pre: BIO 360 and junior standing or permission of the instructor.

BIO 425G

Marine Biodiversity: A Larval Perspective

(3 crs.) Almost all marine life has a biphasic life cycle. This course explores the diversity marine reproduction and larval forms, and how humans interact with species from a larval perspective. (Lec. 3) Pre: BIO 262 and junior standing, or permission of instructor. (D1) (B2) (GC)

BIO 437

Fundamentals of Molecular Biology

(3 crs.) Cross-listed as (BIO), CMB 437. Biochemical basis of heredity as seen through the structure and function of nucleic acids. Includes DNA replication, transcription, translation, gene regulation, and gene organization in prokaryotes and eukaryotes. Current methods emphasized. (Lec. 3) Pre: CMB 211, BIO 352, and CMB 311, or permission of instructor.

BIO 439

Big Data Analysis

(3 crs.) Cross-listed as (BIO), DSP 439. Learn about big data and how to write scripts to analyze data. (Lec. 3) Pre: junior standing, MTH 131 or 141. Not for graduate credit.

BIO 440G

How Our Genes and the Environment Shape Our Lives

(3 crs.) Explore how deteriorating environments alter early human embryological development, increasing diseases and neurological disorders, altering entire societies. (Lec. 3) Pre: BIO101 and junior standing. (A1) (B2) (GC)

BIO 444

Insect Ecology

(3 crs.) Ecology of insects and other terrestrial arthropods at the physiological, individual, population, community, and ecosystem levels. Not for graduate credit. (Lec. 3) Pre: BIO 262 or ENT 385.

BIO 452

Advanced Topics In Genetics

(3 crs.) Cross-listed as (CMB), BIO 452. More detailed treatment of topics introduced in the general genetics course (352) including aspects of transmission genetics, molecular genetics, cytogenetics, biotechnology, developmental genetics, and the impact of genetics on society. (Lec. 3) Pre: BIO 352.

BIO 455

Marine Ecology

(3 crs.) Investigation of the structure and dynamics of various marine ecosystems. Includes mineral cycling, energy flow, community and population organization, and behavioral ecology in selected marine environments. (Lec. 3) Pre: 262 or permission of instructor.

BIO 457

Marine Ecology Laboratory

(1 cr.) Field and laboratory work on community relationships of dominant organisms in Rhode Island marine environments. (Lab. 3) Pre: concurrent enrollment in 455. Limited to 15 students.

BIO 465

Coral Reef Ecology

(3 crs.) An introduction to coral reefs, the processes generating their exuberant biodiversity, the effects of human activities on them and the promising ways to restore and preserve these unique ecosystems. (Lec. 3) Pre: BIO 262 or permission of instructor.

BIO 465X

Coral Reef Ecology

(3 crs.) Examine the processes that generate the biodiversity on coral reefs, evaluate the impacts of human activities on them and recognize the strategies to preserve these unique ecosystems. (Lec 3.) Pre: BIO 262, BIO 360 or permission of the instructor. Not open to students with credit in BIO 475.

BIO 467

Animal Behavior

(3 crs.) Roles of natural selection, individual learning, and cultural transmission in shaping animal behavior. (Lec. 3) Pre: two semesters of biology.

BIO 469

Tropical Marine Invertebrates

(5 crs.) Systematic survey of tropical invertebrates. Emphasis on examples from Bermuda's marine environment. Laboratory includes field collections, identification, and preparatory techniques for taxonomic studies. (Practicum, Lab. 8) Taught in Bermuda. Pre: BIO 360, junior standing, and permission of instructor.

BIO 472

Advanced Evolutionary Biology

(3 crs.) Cross-listed as (BIO), GEO 472. A survey of modern evolutionary biology, including classic evolutionary theory, phylogenetics, evolution and development, adaptation, mass extinction and genomic evolution. (Lec./Sem. 3) Pre: BIO/GEO 272, or permission of instructor. Not for graduate credit.

BIO 475

Ecology of Coral Reefs in Bermuda

(5 crs.) Structure and function of coral reef ecosystems with emphasis on the biology of corals. Laboratory sessions focus on field surveys and research techniques. (Practicum, Lab 8) Taught in Bermuda. Pre: BIO 262 and junior standing; SCUBA certification required.

BIO 480

Community Ecology

(3 crs.) Exploration of community ecology, with an emphasis on interspecific interactions (competition, predation, mutualism), species diversity, succession, niche theory, and island bio-geography. Format includes lecture, case studies, and discussion. (Lec. 3) Pre: BIO 262 or permission of instructor. Not for graduate credit.

BIO 481X

Biology of Human Health and Disease

(3 crs.) Application of biological concepts to understand the basis of human diseases and how biology relates to their treatment. Topics include traditional medicine, medical genetics, cancer, and molecular medicine. (Lec. 2, Seminar 1) Pre: BIO 101, BIO/CMB 352 and one or more additional BIO courses at 200 level or above. Not for graduate credit.

BIO 482G

Biology of Human Health and Disease

(3 crs.) An applied study of the evolutionary drivers of human health and of our current understanding of the ecological, genetic, and molecular mechanisms underpinning disease conditions. (Lec. 3) Pre: BIO/CMB 352 and one additional BIO course at 200 level or above. Not for graduate credit. (D1) (GC)

BIO 485

Salt Marsh Ecology

(4 crs.) Cross-listed as (BIO), NRS 485. Structure and function of salt marsh ecosystems, including biogeochemistry, in the context of global change. Seminar-style discussions of primary scientific literature, laboratory and field exercises, and an independent research poster. (Lec. 2, Lab 4) Pre: BIO 262 or NRS 223 and 2 semesters of chemistry or permission of instructor. BIO 360 recommended. Not for graduate credit.

BIO 491

Independent Biological Research

(1-3 crs. each) Individualized laboratory, field, or literature research projects. May be repeated for a total of 6 credits. (Independent Study) Pre: open only to undergraduates on arrangement with staff. S/U only.

BIO 492

Independent Biological Research

(1-3 crs. each) Individualized laboratory, field, or literature research projects. May be repeated for a total of 6 credits. (Independent Study) Pre: open only to undergraduates on arrangement with staff. S/U only.

BIO 495

Tropical Marine Biology Research

(6 crs.) Independent marine research in Bermuda. Topics may include marine ecology, physiology, systematics, etc. Proposal, oral report, and project paper required. (Practicum, Lab 12). Taught in Bermuda. Pre: Junior standing, BIO 475 and 469.

BIO 498

Teaching Practicum in Biological Sciences

(1 cr.) Teaching experience for undergraduates through planning and assisting in introductory laboratory courses. Can be repeated once for credit. Not for major credit in BS BIO, BA BIO, or BS MBIO. (Prac. 1). Pre: permission of instructor. Not for graduate credit.

BIO 502

Introduction to Neurobiology

(4 crs.) Cross-listed as (BIO), NEU 502. Fundamental processes in neurobiology with emphasis on cellular and membrane mechanisms of nerve functioning. (Lec 3). Pre: BIO 201 and MTH 141, or permission of instructor.

BIO 508

Seminar in Biological Literature

(1 cr.) Cross-listed as (BIO), CMB, AFS, AVS, PLS, NRS, LIB 508. Survey of biological literature including traditional methods of bibliographic control, contemporary information retrieval services, and the development of a personalized information system. (Sem. 1) Pre: graduate standing or permission of the instructor.

BIO 511

Special Readings in Developmental Plant Anatomy

(3 crs.) Intensive tutorial work, research, and reading on ontogeny of plant structures and morphogenetic mechanisms. (Independent Study) Pre: graduate standing and permission of instructor. Concurrent audit of 311 required. Offered on demand.

BIO 512

Evolution and Diversity of Fishes

(4 crs.) Origin, evolution, and diversification of fishes, their phylogenetic relationships, and morphological, physiological, ecological, and behavioral adaptations in marine and freshwater habitats. (Lec. 3, Lab. 3) Pre: Graduate standing and permission of instructor.

BIO 517

Herpetology

(4 crs.) Cross-listed as (NRS), BIO 517. This course provides an in-depth background on the biology, ecology, conservation, and management of reptiles and amphibians, including field methods for studying amphibians and reptiles of northeastern North America. (Lec. 2, Lab. 4) Pre: graduate student in biological and environmental sciences and permission of instructor.

BIO 519

Field Experience in Herpetology

(1 cr.) Cross-listed as (NRS), BIO 519. Capstone field trip in herpetology to region with higher amphibian and reptile diversity, such as Appalachia, to hone skills in identification, broaden understanding of ecology, and apply field research methods. (Practicum) Pre: Concurrent enrollment or credit for NRS/BIO 517 and permission of instructor. S/U only.

BIO 522

Sensory Ecology

(3 crs.) Cross-listed as (BIO), NEU 522. Sensory system structure and function and the opportunities and constraints presented by the sensory environment. Lectures on fundamentals and discussion of case studies and problems in the primary literature. (Lec. 1, Seminar 2) Pre: Graduate standing or permission of instructor.

BIO 539

Big Data Analysis

(3 crs.) Cross-listed as (BIO), DSP 539. Learn about big data and gain sufficient programming skills to analyze data efficiently and accurately for research. (Lec. 3) Pre: graduate standing

BIO 541

Comparative Physiology of Marine Animals

(3 crs.) Comparison of physiological mechanisms by which animals maintain life with emphasis on marine invertebrates. Responses to external environment mediated by receptors, nervous systems, effectors. Living control systems for muscular activity and circulation. (Lec. 3) Pre: one physiology course. In alternate years.

BIO 544

Insect Ecology

(3 crs.) Cross-listed as (ENT), BIO 544. Ecology of insects and other terrestrial arthropods at the physiological, individual, population, community, and ecosystem levels. Pre: permission of instructor. In alternate years.

BIO 550

Advanced Topics In Neurobiology

(3 crs.) Published papers in selected aspects of neurobiology will be discussed. Representative topics include role of Ca++, c-AMP in the nervous system, gating currents learning at the cellular level, cellular rhythmicity. (Seminar) In alternate years.

BIO 563

Biology and Ecology of Fishes

(4 crs.) Cross-listed as (BIO), NRS 563. Exploration of the functional biology and ecology of marine and freshwater fishes through lecture and discussion of primary literature. Laboratory involves specimen study, field trips, and a research project. (Lec. 3, Lab. 3) Pre: BIO 366 or equivalent, or permission of instructor.

BIO 572

Advanced Evolutionary Biology

(3 crs.) Cross-listed as (BIO), GEO 572. A survey of modern evolutionary biology, including classic evolutionary theory, phylogenetics, evolution and development, adaptation, mass extinction and genomic evolution. (Lec./Sem. 3) Pre: BIO/GEO 272, graduate standing, or permission of instructor.

BIO 579

Advanced Genetics Seminar

(1 cr.) Cross-listed as (CMB), BIO 579. Current topics in genetics, including cytological, ecological, molecular, physiological, population, quantitative, and radiation genetics. (Seminar) Pre: CMB 352 and permission of instructor.

BIO 580

Community Ecology

(3 crs.) Explores community ecology, with an emphasis on interspecific interactions (competition, predation, mutualism), species diversity, succession, niche theory, and island biogeography. Format includes lecture, case studies, and discussion. (Lec. 3) Pre: BIO 262 or permission of instructor.

BIO 585

Salt Marsh Ecology

(4 crs.) Cross-listed as (BIO), NRS 585. Structure and function of salt marsh ecosystems, including biogeochemistry, in the context of global change. Seminar-style discussions, weekly assignments, written and oral presentations of independent proposal and research project. (Lec. 2, Lab. 4) Pre: permission of instructor. Prior undergraduate coursework equivalent to a course in ecology and 2 semesters of chemistry is expected.

BIO 586

Medical and Veterinary Entomology

(3 crs.) Cross-listed as (BIO), ENT 586. Life history, classification, habits, and control of insects and other arthropods affecting human and animal health. Topics will include public health significance, vector-parasite interactions, and survey and research methodologies. (Lec. 3, Lab. 4) Pre: ENT 331 or 381 or equivalent. In alternate years.

BIO 587

Seminar In Neurobiology

(1 cr.) Cross-listed as (BIO), NEU 587. Survey of current literature in the neurosciences. Topics include molecular and behavioral electrophysiology, ion channels, nerve net modeling, ultrastructure of excitable cells, receptor and pharmacological neurobiology of invertebrates and vertebrates. (Seminar) Pre: graduate standing or one advanced neuroscience course.

BIO 591

Independent Biological Research

(1-6 crs.) Individualized laboratory, field, or literature research projects. May be repeated for a total of 6 credits. Pre: graduate standing and permission of instructor. S/U credit.

BIO 592

Independent Biological Research

(1-6 crs.) Individualized laboratory, field, or literature research projects. May be repeated for a total of 6 credits. Pre: Graduate standing and permission of instructor. S/U credit.

BIO 593

Special Topics in Biological Sciences

Selected areas pertinent to needs of individuals or small groups. Class, seminar or tutorial. Topics may include the following: biomechanics, cell biology, ecological morphology, functional morphology, ichthyology, molecular biology, morphology and mechanics, physiology, plant cell development and zoology. May be repeated for a total of 6 credits.

BIO 594

Special Topics in Biological Sciences

Selected areas pertinent to needs of individuals or small groups. Class, seminar or tutorial. Topics may include the following: biomechanics, cell biology, ecological morphology, functional morphology, ichthyology, molecular biology, morphology and mechanics, physiology, plant cell development and zoology. May be repeated for a total of 6 credits.

BIO 599

Master's Thesis Research

(1-9 crs.) Number of credits is determined each semester in consultation with the major professor or program committee. (Independent Study) S/U credit.

BIO 642

Seminar In Physiology

(1-3 crs.) Reports and discussions on topics of current research in physiology. Subject matter adapted to meet interests of students. (Seminar) Pre: permission of instructor.

BIO 654

Seminar in Ichthyology

(2 crs.) Reading, library research, reports, and class discussion on problems of current research interest in the biology of fishes. (Seminar) Pre: BIO 563 or permission of instructor. In alternate years.

BIO 675

Advanced Ecology Seminars

(2 crs.) Specialized and advanced areas of ecological research and theory, including biogeography, Pleistocene ecology, population dynamics, energy flow in ecosystems, and radiation ecology. (Seminar) Pre: permission of instructor.

BIO 691

Biological Problems

(1-6 crs.) Special work to meet the needs of individual students who are prepared to undertake special problems. (Independent Study) Pre: permission of chairperson. Open only to doctoral students.

BIO 692

Biological Problems

(1-6 crs.) Special work to meet the needs of individual students who are prepared to undertake special problems. (Independent Study) Pre: permission of chairperson. Open only to doctoral students.

BIO 699

Doctoral Dissertation Research

(1-12 crs.) Number of credits is determined each semester in consultation with the major professor or program committee. (Independent Study) S/U credit.